pyapksigner
pyapksigner can sign the apk file.
$ pip install pyapksigner
$ pyapksigner --sign [apk_path]
> [signed_apk_path]
$ pyapksigner --sign --key_path="sample.jks" --key_alias="sample" --key_pass="sample_key" --ks_pass="sample_ks"
> [signed_apk_path]